I can't seem to find a 开发者_StackOverflow社区command for it under Options->Keyboard.
You can assign shortcuts to Resharper.ReShaper_Dfa_ReachingDefinitionsUp and Resharper.ReShaper_Dfa_ReachingDefinitionsDown commands for 'Value Origin' and 'Value Destination' respectively.
Try typing reachingdefinition
as a search string in the VS shortcuts window.
The closest I could find was using the default keyboard shortcut (for R#2.x/IntelliJ) of Ctrl+Alt+Shift+A for 'Inspect this'. This pops up a context menu with entries 'Value Origin' and 'Value Destination'. Not a direct keyboard shortcut, but at least you don't have to take your hands off the keyboard.
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